Trigonochlamydidae

Trigonochlamydidae; Temporal range: Recent[1]
	Ghost slug, Selenochlamys ysbryda
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
(unranked):	clade Heterobranchia; clade Euthyneura; clade Panpulmonata; clade Eupulmonata; clade Stylommatophora; informal group Sigmurethra; clade limacoid clade
Superfamily:	Parmacelloidea
Family:	Trigonochlamydidae; Hesse, 1882
Diversity
	9 genera, 11 species

Trigonochlamydidae is a family of air-breathing land slugs, terrestrial gastropod molluscs in the clade Eupulmonata (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).

Distribution

The native distribution of Trigonochlamydidae includes Caucasus,[1] Iran and Turkey.[2]

Taxonomy

Previously, Trigonochlamydidae was placed in the superfamily Trigonochlamydoidea Hesse, 1882, in the subinfraorder Sigmurethra. This was the only family within that superfamily.[3]

The family Trigonochlamydidae is now classified within the limacoid clade, which itself belongs to the clade Stylommatophora, within the clade Eupulmonata (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).

The following two subfamilies have been recognized in the taxonomy of Bouchet & Rocroi (2005):

subfamily Trigonochlamydinae Hesse, 1882 - synonyms: Selenochlamydinae I. M. Likharev & Wiktor, 1980
subfamily Parmacellillinae Hesse, 1926[4]

Genera

There are nine[2] genera in the family Trigonochlamydidae with a total of 11 species:

Subfamily Trigonochlamydinae

Boreolestes Schileyko & Kijashko, 1999
- Boreolestes likharevi Schileyko & Kijashko, 1999 - type species[5]
- Boreolestes sylvestris Kijashko in Schileyko & Kijashko, 1999[5]
Drilolestes Lindholm, 1925 - with only one species Drilolestes retowskii (O. Boettger, 1884)[5]
Hyrcanolestes Simroth, 1901 - with only one species Hyrcanolestes velitaris (Martens, 1880)[5]
Khostalestes Suvorov, 2003[2] - with only one species Khostalestes kochetkovi Suvorov, 2003[2][5]
Lesticulus Schileyko, 1988 - with only one species Lesticulus nocturnus Schileyko, 1988[5]
Trigonochlamys O. Boettger, 1881 - type genus of the family Trigonochlamydidae, with only one species Trigonochlamys imitatrix O. Boettger, 1881[5]
Troglolestes Ljovushkin & Matiokin, 1965 - with only one species Troglolestes sokolovi Liovushkin & Matiokin, 1965[5]
Selenochlamys O. Boettger, 1883
- Selenochlamys pallida Boettger, 1883 - type species
- Selenochlamys ysbryda Rowson & Symondson, 2008[6]

Subfamily Parmacellillinae

Parmacellilla Simroth, 1910 - type genus of the subfamily Parmacellillinae, with only one species Parmacellilla filipowitschi Simroth, 1910[7]

Synonym:

Pseudomilax O. Boettger, 1881 is a synonym for various genera in Trigonochlamydidae.

Cladogram

The following cladogram shows the phylogenic relationships of this family with the other families in the limacoid clade:[8]

limacoid clade

Staffordioidea

Dyakioidea

Gastrodontoidea

	Euconulidae

	Trochomorphidae

	Gastrodontidae

	Oxychilidae

Parmacelloidea

	Trigonochlamydidae

	Parmacellidae

	Milacidae

Zonitoidea

Zonitidae

Helicarionoidea

Helicarionidae

	Ariophantidae

	Urocyclidae

Limacoidea

Vitrinidae

Boettgerillidae

	Limacidae

	Agriolimacidae

References

"Family summary for Trigonochlamydidae". AnimalBase, last modified 26 August 2005, accessed 1 September 2010.
Suvorov A. N. (2003). "A new species and genus of carnivorous slugs (Pulmonata Trigonochlamydidae) from West Transcaucasia". Ruthenica 13: 149-152. abstract.
Sistemica (Spanish list of genera), cited 7 October 2008.
Hesse P. (1926). "Die Nacktschnecken der palearktischen Region". Abhandlungen des Archiv für Molluskenkunde 2(1): 1-152. page 47 and page 54. plates 1-2.
Kantor Yu I., Vinarski M. V., Schileyko A. A. & Sysoev A. V.(published online on December 22, 2009). "Catalogue of the continental mollusks of Russia and adjacent territories". Version 2.3.
Rowson B. & Symondson O. C. (2008) "Selenochlamys ysbryda sp. nov. from Wales, UK: a Testacella-like slug new to western Europe (Stylommatophora: Trigonochlamydidae)". Journal of Conchology 39(5): 537-552. abstract.
Schileyko A. A. (2003). "Treatise on Recent Terrestrial Pulmonate Molluscs. Part 11. Trigonochlamydidae, Papillodermidae, Vitrinidae, Limacidae, Bielziidae, Agriolimacidae, Boettgerillidae, Camaenidae." Ruthenica, Supplement 2(11): 1467-1626, Moscow. page 1467.
Hausdorf B. (March 2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". Journal of Biogeography 27(2): 379-390. doi:10.1046/j.1365-2699.2000.00403.x, JSTOR.

External links

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[AB-1] "Family summary for Trigonochlamydidae". AnimalBase, last modified 26 August 2005, accessed 1 September 2010.

[Suvorov_2003-2] Suvorov A. N. (2003). "A new species and genus of carnivorous slugs (Pulmonata Trigonochlamydidae) from West Transcaucasia". Ruthenica 13: 149-152. abstract.

[3] Sistemica (Spanish list of genera), cited 7 October 2008.

[4] Hesse P. (1926). "Die Nacktschnecken der palearktischen Region". Abhandlungen des Archiv für Molluskenkunde 2(1): 1-152. page 47 and page 54. plates 1-2.

[Kantor_2009-5] Kantor Yu I., Vinarski M. V., Schileyko A. A. & Sysoev A. V.(published online on December 22, 2009). "Catalogue of the continental mollusks of Russia and adjacent territories". Version 2.3.

[6] Rowson B. & Symondson O. C. (2008) "Selenochlamys ysbryda sp. nov. from Wales, UK: a Testacella-like slug new to western Europe (Stylommatophora: Trigonochlamydidae)". Journal of Conchology 39(5): 537-552. abstract.

[7] Schileyko A. A. (2003). "Treatise on Recent Terrestrial Pulmonate Molluscs. Part 11. Trigonochlamydidae, Papillodermidae, Vitrinidae, Limacidae, Bielziidae, Agriolimacidae, Boettgerillidae, Camaenidae." Ruthenica, Supplement 2(11): 1467-1626, Moscow. page 1467.

[8] Hausdorf B. (March 2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". Journal of Biogeography 27(2): 379-390. doi:10.1046/j.1365-2699.2000.00403.x, JSTOR.