Neotheropoda

Neotheropoda (meaning "new theropods") is a clade that includes coelophysoids and more advanced theropod dinosaurs, and is the only group of theropods that survived the Triassic–Jurassic extinction event. All neotheropods became extinct by the end of the Early Jurassic period except for Averostra.

Neotheropods
Temporal range:
Late Triassic–Present,
Cast of a Coelophysis bauri skeleton
A female house sparrow (Passer domesticus)
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Clade: Neotheropoda
Bakker, 1986
Subgroups
Synonyms
  • Avepoda Paul, 2002
  • Eutheropoda Novas, Haro & Canale, 2003

Classification

Neotheropoda was named by Robert T. Bakker in 1986 as a group including the relatively derived theropod subgroups Ceratosauria and Tetanurae, and excluding coelophysoids.[1] However, most later researchers have used it to denote a broader group. Neotheropoda was first defined as a clade by Paul Sereno in 1998 as Coelophysis plus modern birds, which includes almost all theropods except the most primitive species.[2] Dilophosauridae was formerly considered a small clade within Neotheropoda, but was later considered to be paraphyletic.[3]

The following family tree illustrates a synthesis of the relationships of the early theropod groups compiled by Hendrickx et al. in 2015.[4]

Theropoda
 

Herrerasauridae

 
 
 

Eoraptor

 
 
 

Eodromaeus

 
 
 

Tawa

 
 
 

Daemonosaurus

 
Neotheropoda
 

Coelophysoidea

 
 
 

Liliensternus

 
 
 

Zupaysaurus

 
Averostra
 

Ceratosauria

 
 

Tetanurae

 
 
 
 
 
 
 
 
 
 

This second cladogram is based on the redescription of Dilophosaurus by Marsh and Rowe (2020).[3]

Theropoda
 
 

Staurikosaurus

 
 

Herrerasaurus

 
 
 
 
 

Tawa

 
 

Chindesaurus

 
 
 
 

Eodromaeus

 
Neotheropoda
 
 

Lepidus

 
 

Liliensternus

 
 
 

Coelophysoidea

 
 
 

Zupaysaurus

 
 
 

Cryolophosaurus

 
 
 

Dilophosaurus

 
 

Averostra

 
 
 
 
 
 
 
 

This third cladogram is based on the larger analysis in the description of Pendraig milnerae by Spiekman et al. 2021:[5]

Theropoda
 

Eodromaeus

 
 
 

Lepidus

 
Neotheropoda
Coelophysoidea
 

Panguraptor

 
 
 

Pendraig

 
 
 

Powellvenator

 
 
 

Lucianovenator

 
 
 

Syntarsus kayentakatae

 
 

Coelophysidae

 
 
 
 
 
 
 
 
 

Liliensternus

 
 

Dracoraptor

 
 
 
 

Zupaysaurus

 
 
 

Gojirasaurus

 
 
 

Cryolophosaurus

 
 
 

Dilophosaurus

 
 
 

Sarcosaurus

 
 
 

Tachiraptor

 
 

Averostra

 
 
 
 
 
 
 
 
 
 
 

References

  1. Bakker, R.T. 1986. The Dinosaur Heresies. William Morrow, New York. 481 pp.
  2. Sereno, 1998. A rationale for phylogenetic definitions, with application to the higher-level taxonomy of Dinosauria. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. 210, 41-83.
  3. Marsh, Adam D.; Rowe, Timothy B. (2020). "A comprehensive anatomical and phylogenetic evaluation of Dilophosaurus wetherilli (Dinosauria, Theropoda) with descriptions of new specimens from the Kayenta Formation of northern Arizona". Journal of Paleontology. doi:10.1017/jpa.2020.14.
  4. Hendrickx, C., Hartman, S.A., & Mateus, O. (2015). An Overview of Non- Avian Theropod Discoveries and Classification. PalArch’s Journal of Vertebrate Palaeontology, 12(1): 1-73.
  5. Spiekman SN, Ezcurra MD, Butler RJ, Fraser NC, Maidment SC (2021). "Pendraig milnerae, a new small-sized coelophysoid theropod from the Late Triassic of Wales". Royal Society Open Science. 8 (10): Article ID 210915. doi:10.1098/rsos.210915.
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