Songlingornithidae

Songlingornithidae is a family of basal euornitheans from the Early Cretaceous of China. All known specimens come from the Jiufotang Formation, dating to the early Aptian age, 120 million years ago.

Songlingornithids
Temporal range: Early Cretaceous,
Holotype of Yixianornis grabaui, Paleozoological Museum of China
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Clade: Avialae
Clade: †Yanornithiformes
Family: †Songlingornithidae
Hou, 1997
Type species
†Songlingornis linghensis
Hou, 1997
Genera
Synonyms
  • Yanornithidae? Zhou & Zhang, 2001
  • Aberratiodontuidae? Gong, Hou & Wang, 2004
  • Yixianornithidae? Zhang & Zhou, 2006

The family Songlingornithidae was first named by Hou in 1997 to contain the type genus, Songlingornis.[1] Clarke et al. (2006) was the first to find a close relationship between Songlingornis and the "yanornithids".[2]

The name was originally coined to reflect a close relationship between the two supposedly similar (but poorly preserved) genera Songlingornis and Chaoyangia. However, subsequent studies found that Chaoyangia was probably not closely related to Songlingornis. Instead, Songlingornis was found to be closely related to another group which had been given the name Yanornithidae—the name Songlingornithidae, having been named first, took precedence.[3] Some later studies have also placed the hongshanornithids in this group.[3][4] At least one study has found the Late Cretaceous Mongolian avialan Hollanda to be a member of the group as well.[5]

Phylogeny

Below is a cladogram from Mayr (2017) showing the relationships between Songlingornithidae and other ornithuromorphs:[6]

Ornithuromorpha
 

Archaeorhynchus

 
 
 

Vorona

 
 
 

Schizooura

 
 
 

Hongshanornithidae

 
 
 

Jianchangornis

 
 
Songlingornithidae
 

Songlingornis

 
 
 

Yixianornis

 
 
 

Piscivoravis

 
 

Yanornis

 
 
 
 
 
 

Gansus

 
 
 

Apsaravis

 
 
 

Hesperornithes

 
 
 

Ichthyornis

 
 

Neornithes (modern birds)

 
 
 
 
 
 
 
 
 
 
 

A 2020 study by Wang et al. failed to recover a monophyletic Songlingornithidae, instead finding Yanornis to form a clade with the newly described genera Abitusavis and Similiyanornis, for which the name Yanornithidae was used:[7]

Ornithuromorpha
 

Archaeorhynchus

 
 
 

Jianchangornis

 
 
 
 

Bellulornis

 
 

Schizoouridae

 
 
 
 

Hongshanornithidae

 
 
 

Songlingornis

 
 
Yanornithidae
 

Abitusavis

 
 

Similiyanornis

 
 

Yanornis

 
 
 
 

Piscivoravis

 
 
 
 

Yixianornis

 
 
 

Apsaravis

 
 
 

Patagopteryx

 
 

Vorona

 
 
 
 
 
 

Iteravis

 
 
 

Gansus

 
 

Ornithurae

 
 
 
 
 
 
 
 
 
 
 

References

  1. Hou, (1997). Mesozoic Birds of China. Taiwan Provincial Feng Huang Ku Bird Park. Taiwan: Nan Tou. 228 pp.
  2. Clarke, Zhou and Zhang, (2006). "Insight into the evolution of avian flight from a new clade of Early Cretaceous ornithurines from China and the morphology of Yixianornis grabaui." Journal of Anatomy, 208: 287-308.
  3. O'Connor, J.K. and Zhou Z. (2012). "A redescription of Chaoyangia beishanensis (Aves) and a comprehensive phylogeny of Mesozoic birds." Journal of Systematic Palaeontology, (advance online publication). doi:10.1080/14772019.2012.690455
  4. Zhou, S.; Zhou, Z.; O'Connor, J. (2013). "A new piscivorous ornithuromorph from the Jehol Biota". Historical Biology: 1. doi:10.1080/08912963.2013.819504.
  5. O’Connor, J. K.; Zhang, Y.; Chiappe, L. M.; Meng, Q.; Quanguo, L.; Di, L. (2013). "A new enantiornithine from the Yixian Formation with the first recognized avian enamel specialization". Journal of Vertebrate Paleontology. 33: 1. doi:10.1080/02724634.2012.719176.
  6. Mayr, G. (2017). "Pectoral girdle morphology of Mesozoic birds and the evolution of the avian supracoracoideus muscle". Journal of Ornithology. 158 (3): 859–867. doi:10.1007/s10336-017-1451-x.
  7. Wang, M.; Li, Z.; Liu, Q.; Zhou, Z. (2020). "Two new Early Cretaceous ornithuromorph birds provide insights into the taxonomy and divergence of Yanornithidae (Aves: Ornithothoraces)". Journal of Systematic Palaeontology. 18 (21): 1805–1827. doi:10.1080/14772019.2020.1836050.


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